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ABSTRACT
We show how a random mutation hill climber that does multi-level selection utilizes transposition to escape local optima on the discrete Hierarchical-If-And-Only-If (HIFF) problem. Although transposition is often deleterious to an individual, we outline two population models where recently transposed individuals can survive. In these models, transposed individuals survive selection through cooperation with other individuals. In the multi-population model, individuals were allowed a maturation stage to realize their potential fitness. In the genetic algorithm model, transposition helped maintain genetic diversity even within small populations. However, the results for transposition on the discrete Hierarchical-Exclusive-Or (HXOR) problem were less positive. Unlike HIFF, HXOR does not benefit from random drift. This led us to hypothesize that two conditions necessary for transposition to enhance evolvability are (i) the presence of local optima and (ii) susceptibility to random drift. This hypothesis is supported by further experiments. The findings of this paper suggest that epistasis and large mutations can sustain artificial evolution in the long-term by providing a way for individuals and populations to escape evolutionary dead ends. Paradoxically, epistasis creates local optima and holds a key to its resolution, while deleterious mutations such as transposition enhance evolvability. However, not all large mutations are equal. REFERENCES
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